Set two oven racks in the upper and lower third. Preheat the oven to 325°F and line two baking sheets with parchment paper.
Lightly crush the potato chips with your hands to about pea-size pieces. Measure 2 cups for the dough and reserve 1/2 cup for topping.
In a medium bowl, whisk together the flour, cornstarch, baking soda, and fine salt until evenly combined.
In a large bowl, beat the softened butter with both sugars on medium speed until pale and fluffy, 2 to 3 minutes. Beat in the vanilla, then the egg, followed by the yolk, mixing just until glossy and smooth.
Mix in about half of the dry ingredients on low speed. Add the chocolate chips and mix briefly to distribute. Add the remaining dry ingredients and mix just until a few flour streaks remain.
Fold in the 2 cups of crushed potato chips with a spatula until no dry spots remain. Let the dough rest 15 minutes (or chill 20 minutes) to hydrate and reduce spreading.
Scoop the dough into 1½-tablespoon mounds and arrange 2 inches apart on the prepared sheets. Gently press the tops into the reserved crushed chips and sprinkle each with a tiny pinch of flaky sea salt.
Bake both sheets at once for 12 to 14 minutes, rotating the pans front-to-back and top-to-bottom halfway through, until edges are golden and centers look set but still pale.
Cool cookies on the baking sheets for 5 minutes, then transfer to a wire rack to cool completely.
Baker’s tip: Spoon-and-level (or weigh) your flour for best texture; too much flour makes cookies dry and cakey.